Eren Kruger and Reiner Braun

After the Battle of Utgard Castle, he fought the Armored Titan transformed by Reiner Braun. Just as he was about to win, he was struck by the Colossal Titan transformed by Bertholdt Hoover and knocked unconscious. He was then taken away by the two, along with Ymir. At Reiner's urging, Ymir joined the trio returning home and took Christa Lance with her. She was later rescued from Reiner by the Survey Corps, who arrived shortly after, but she and Mikasa were both in danger.

Reiner is kicked awake by Annie. Upon waking, he sees Armin Arlert, Connie Springer, Jean, Gabi, Annie, Falco, and others. They then join Hange, Levi, Magath, Pieck, and others. The group decides to put aside their past grievances and work together to stop the Rumbling. Around the campfire, Reiner confesses to having caused Marco's death, and is brutally beaten by Jean.

The following day, the group decided to first rescue the airship technicians from the Yeagerists on the island, and then use the aircraft to approach the Founding Titan. Their first encounter was with the Yeagerists led by Floch Forster. During the battle, Reiner and Annie transformed into Titans and clashed with the Yeagerists soldiers, covering the airship technicians' retreat. Although they successfully protected their teammates, the Armored Titan and the Female Titan were also fiercely attacked by the Yeagerists's Thunder Spear and nearly killed. Finally, Falco transformed into a Titan and managed to subdue the enemy, allowing the group to escape by ship.

They then learned that the Rumbling Titans had landed in Marley and that Liberio had been crushed. Exhausted, Annie and Reiner said their final farewells and then withdrew with Gabi and Falco. Due to damage to the plane and the imminent arrival of the Rumbling Titans, Hange sacrificed herself to buy time, and the plane finally took off.

On the plane, Eren told everyone in his mental world that there was no need to talk, and that they should try to stop him if they wanted, thus completely eliminating any possibility of the group negotiating with Eren.

The plane finally reached directly above the Founding Titan, but was shot down by the Beast Titan summoned by the Founding Titan. The group forced their way onto the Founding Titan's back. Reiner transformed into the Armored Titan and defeated the Beast Titan, only to discover it was just an empty shell. Armin was Armin Arlert captured by the Founding Titan, who simultaneously summoned the Nine Titans from different eras to fight Reiner and the others. The group was quickly overwhelmed, Armored Titan's head was bitten off by the Colossal Titan summoned by the Founding Titan.

Reiner had lost count of how many times he had been defeated, only managing to avoid falling thanks to Jean's grab. Reiner declared he could transform into a Titan one more time, to which Jean retorted, "Who has your broken Titan ever beaten except for the door? Even though you're the Armored Titan, your Armored Titan always gets shattered." In their dire straits, the group was rescued by Falco, Annie, Gabi, and others who arrived just in time. It turned out that Falco gained the ability to fly after transforming into a Titan. After being rescued by Falco's Jaw Titan, Levi rewrote the plan, changing the mission to drawing the attention of the War Hammer Titan to buy Jean time to blow up Eren's Titan-transformed neck. Reiner engaged the War Hammer Titan summoned by the Founding Titan, thus providing cover for Jean.

Annie and the others joined the battle. Reiner single-handedly held off all the War Hammer Titan summoned by Eren, but he was also severely injured. Just as the group was in despair, some of the Nine Titans from previous generations began to help them, such as Bertolt's Colossal Titan and the Gariad brothers' Jaw Titan. At the same time, Zeke volunteered to be killed by Captain Levi, Rumbling was temporarily stopped. Reiner used explosives to blow off the head of Eren 'Founding Titan, and the source of the Titan's power— the Hallucigenia —rushed out, attempting to reconnect Eren's head. At this moment, Reiner rushed out and firmly controlled the Hallucigenia. Meanwhile, Armin Arlert was rescued, transformed, and launched an explosion, shattering Eren's Skeleton Titan into pieces.

Eren's Founding Titan was destroyed, and the group thought it was finally over. However, the Hallucigenia not dead. It spewed gas, instantly transforming all the Eldians near the military base into Pure Titans to protect Hallucigenia. Reiner was shocked to discover that Jean, Connie Springer, Gabi, and his mother had all been transformed into Pure Titans. Eren then reappeared, this time as the Colossal Titan, and engaged Armin Arlert's Colossal Titan in battle. Reiner, Annie, Pieck, faced with the reality that their loved ones had all been transformed into Pure Titans, tearfully continued their attempt to kill the Hallucigenia.

In the end, Mikasa made her decision. With the help of Armin Arlert and Levi, she charged into Eren's Colossal Titan mouth and severed Eren's head.

In chapter 139, due to Eren's death, the power of the Titans dissipates, and Reiner and Karina, who has returned to being human, are reunited.

Bertholdt Hoover, a member of the Eldian Giants Warrior Unit of the Marleyan Imperial Military. He was a key figure in the operations on Paradis Island, a fellow Marley of Reiner Braun, Annie Leonard, and Ymir Beli. His superior was Marley Army General Dio Magath.

He infiltrated the walls as an undercover agent for five years, graduating third in his class from the Southern 104th Training Corps before joining the Survey Corps. His long time together forged a strong bond with Reiner. He was gentle but lacked initiative and was not particularly noticeable. He graduated third in his class from the 104th Training Corps and later joined the Survey Corps. He was devoured by Armin, who had transformed into a Titan, during the Battle of Wall Maria. In the War for Heaven and Earth, Armin awakened his consciousness from the Coordinate, helping him fight against Eren.

Reiner and Annie's fellow townsman. Because of his unusual sleeping posture, he is often the target of teasing by his classmates, and his sleeping posture is also used as a source of funny weather predictions.

His birthplace is "Homeland" (i. e., Marley), and according to the records, he, Reiner, and Annie came from a village deep in the mountains southeast of the Wall Maria.

His distinguishing features are tall, dark-haired man with gentle, droopy eyes and a taciturn nature.

He had mastered all the skills and was the most promising of all the soldiers. However, he lacked initiative and often pushed tasks onto others. Reiner commented, "He clearly has outstanding abilities but always follows behind others."

Reiner and was his companion.

Initially, they volunteered to join the Military Police Brigade upon entering the training corps.

In his conversation with Reiner mentioned that Bertholdt had feelings for Annie.

Unlike Reiner, she did not create other personalities to relieve psychological stress, but she always felt guilty about her actions five years ago; her low-key and passive demeanor was also to cover up this contradictory feeling.

His biggest flaws are a lack of clear understanding of what he should do and a lack of initiative. Therefore, he doesn't stand out in the highly individualistic 104th Training Corps and is overshadowed by Eren and Reiner. He himself seems to understand this.

He chose to become a Titan warrior because his father was seriously ill, hoping that his identity would allow his ailing father to receive care from the Marley government.

One of the seven warriors of Marley, the host of the Colossal Titan, the "God of Destruction," possesses exceptional marksmanship and a strong learning ability. Reiner commented, "He mastered the use of the Colossal Titan almost instantly." He participated in the "Founding Titan Recovery Operation" alongside Marcel Galliard Galliad, Reiner Braun, and Annie Reinhardt, forming a strong bond with his teammates.

When Marcel Galliard was lost, he considered ceasing the fighting and returning to Marley, but was stopped by Reiner, and later destroyed the Wall Maria.

He remained deeply troubled by the story of the villager who helped them but later hanged himself out of guilt, and he often pondered it.

The official novel mentions that his unusual sleeping posture stemmed from nightmares.

His true identity is the Colossal Titan, a member of the Marley Warriors. Even as a Warrior candidate, he already possessed excellent marksmanship skills.

He inherited the Colossal Titan in 843 and instantly destroyed a military base during the war, earning him the title of "God of Destruction" from the Marleyan military.

Hannes was eaten by the same Titan who ate his mother years ago, Eren nearly suffered a mental breakdown and fell into despair. Mikasa also had no hope of survival, made a poignant confession to Eren, thanking Eren for everything he had done for her, and smiled happily. Deeply moved, Eren recalled his mother's words, rallied his spirits, and resolved to protect Mikasa, making a pact with the Red Scarf. In the end, Eren unleashed a powerful will to protect Mikasa, and then he gained the power of the "Coordinate" (the ability to seemingly control other Pure Titans with his mind).

The Pure Titans, now under their control, attacked and devoured the Titan who had eaten their mother. Later, due to their hostility towards the Armored Titan, a large number of Pure Titans turned their attacks on the Armored Titan and the Colossal Titan. After escaping the Titans' clutches, Eren carried the injured Mikasa and escaped the danger with his Survey Corps comrades. In reality, Eren was able to activate the "Coordinate" power because he was the host of the Founding Titan, and Dina Fritz, the Titan who ate Eren's mother and Hannes, possessed the King Fritz bloodline. When Founding Titan non-royal host of the Founding Titan comes into contact with a Titan with royal blood, the "Coordinate" power is activated.

Eren and his surviving Survey Corps comrades retreated to Wall Rose. The Allied Forces (main forces of the Survey Corps and the Military Police Brigade) that had been dispatched to rescue Eren numbered around one hundred, but 60% of them perished during the retreat, leaving Eren deeply remorseful. During Eren's period of confusion, Jean Kirstein used his "Eren- style preaching" to help Eren re-establish his goals: to use his "Coordinate" ability to command Titans to block Wall Maria and to personally kill Reiner and others. A week later, after recovering from his injuries, Levi, after listening to Hange Zoe and Connie Springer's information about Titans, discussed with Erwin Smith, who was also recovering, and decided to form a new Levi Squad. Eren, along with Mikasa, Armin, and other comrades from the 104th Training Corps, joined the new Levi Squad.

During the first internal conflict on Reiner Paradis Island, Eren and the Survey Corps temporarily hid in a cabin in the mountains. Under Hange and Levi's guidance, they conducted research and experiments to explore the characteristics of Titan transformation. They obtained information about the limit of Eren's Titan transformations and the level of intelligence he gained after transforming. However, they were disappointed to learn that Eren still had no idea how to Hardening, making Armin Arlert's plan to "use the Hardening remains of the Titans to seal the city gates" impossible. Meanwhile, their classmate Krista Lenz revealed her identity to everyone: her real name was Historia Reiss, the illegitimate daughter of the Reiss Family, a true royal family within the walls, who disguised themselves as ordinary nobles.

At this moment Hange and Levi receive a report from Erwin in the central command that someone there intends to kidnap Eren and Historia. Despite Hange and the others' best efforts to protect them, Eren and Historia fall into the hands of the Interior Military Police. In the secret underground cavern of the Reiss Family manor, Eren is chained to a high platform, while Historia is reunited with her biological father, Road Reiss. Through physical contact with the Reiss father and daughter, some of Eren's dormant memories are awakened: it turns out that Eren's father, Grisha Yeager, was the wielder of the Titan power. On the night Wall Maria fell, Grisha Yeager came to the Reiss Family's cellar, defeated and devoured the Titan that Road's eldest daughter, Frieda Reiss, had transformed into, thus obtaining the Reiss Family's inherited power, the "Founding Titan," which could control all Titans. He also killed all of Road's other children present.

Grisha Yeager later found Eren in the refugee camp, took Eren to the woods, left Eren the key to a secret basement, and injected Eren with a Titan-transforming agent. Grisha was then devoured by the Titan-transformed Eren, allowing him to transfer his two Titan powers and memories into Eren's body. Eren was Eren taken back to the refugee camp by Keith Shadis, the training corps' chief instructor. Upon learning the truth, Eren suffered a mental breakdown, believing that he and his father were the root of all the disasters. He cried out, demanding that Historia eat him to reclaim the Founding Titan, and with his royal blood, unleash the Founding Titan's power to control the Titans and end the war.

Historia realized that although the Reiss Family had controlled the Titans for over a hundred years, they had done nothing to save humanity. She refused her father's request to consume Eren's Titan power, smashed the syringe, and attempted to free Eren. At this moment, the Survey Corps arrived to rescue the two. Desperate, Road licked the Titan-transforming syringe that had spilled on the ground, transforming into a maimed, crawling Colossal Titan, which also damaged the cellar, bringing it to the brink of collapse.

Just as the entire legion was about to perish in the collapsing cave, Eren discovered a bottle labeled "Armored Titan" that Road had brought along with the Titan transformation injection. In a desperate move, he bit through the bottle and transformed into a Titan, successfully Hardening his entire body to support the cave and save everyone.

Afterwards, Eren and his army headed to the Orvud District to stop the approaching Titan, Road. Eren transformed into a Titan and threw a large amount of explosives into Road's body, shattering it. Finally, Historia personally delivered the final blow to her father, thus establishing the authority of the new queen who would soon ascend the throne.

With the new queen's ascension to the throne, the Survey Corps eliminated the uncertainties surrounding her. Eren mastered the ability to Hardening and used it to complete Hange's design for the "Titan Guillotine," testing it in Trost District and killing a large number of Titans. All of this laid the foundation for the next step in the plan to reclaim human territory— Wall Maria.

Meanwhile, through increased contact with Historia, Eren awakened more fragments of memories passed down from his father. From these memories, Eren discovered that Keith Shadis, a Training Corps instructor and former Survey Corps commander, was acquainted with Grisha Yeager from the past and might know some of Grisha Yeager's secrets. Eren and the others visited Keith. However, Keith's knowledge of Grisha Yeager's origins was extremely limited. But Eren learned from Keith about his mother Carla Yeager's expectations for him: even if one is ordinary, simply being born into this world is already a great achievement. This relieved Eren's obsession with being "not special."

Subsequently, Operation Battle to Retake Wall Maria. Eren and his army rushed to the Shiganshina district overnight, and successfully sealed the breach on the outside of Shiganshina as Fudō, using their Titan transformation and Hardening abilities. However, the fact that Reiner and Bertolt remained inactive and whose whereabouts were unknown made everyone very uneasy. Armin Arlert realized that the enemy might be hiding inside the wall and successfully forced Reiner out.

On the other side of the wall, the Warrior Commander, who came from Reiner and the others' "hometown," also transformed into the Beast Titan, and together with a large number of Pure Titans under his control, blocked the Survey Corps' retreat. Eren, Reiner, Erwin, Levi, and the Warrior Commander engaged in a battle of wits inside and outside Wall Maria. Inside the Shiganshina District, Eren unleashed his long-trained Hardening —concentrating Hardening on his fists to greatly increase their hardness—easily shattering Reiner's full-body Armored Titan. Combined with the Survey Corps' new weapon— the Thunder Spear —he quickly cornered Reiner.

At this moment, Bertolt appeared and transformed into the Colossal Titan. Faced with its unparalleled destructive power, the situation instantly reversed. Armin Arlert sacrificed himself to launch a feint attack to attract Bertolt's attention, while Eren left behind his Hardening Titan body. After his main body separated, he went straight for Bertolt's main body, which was protruding from the back of the Colossal Titan's neck, and successfully captured him alive.

However, Armin Arlert suffered severe burns from the intense heat of the steam during the feint attack, leaving him on the verge of death. Meanwhile, Commander Erwin, leading his troops as bait, was also critically wounded and near death by the Beast Titan. The last vial of Titan-transforming serum in Captain Levi's possession was crucial in determining who would survive. Driven mad, Eren desperately begged Levi to save Armin Arlert. After a moment of inner turmoil, Levi decided to release Erwin from this world and administered the serum to Armin Arlert.

Armin Arlert, now a Titan, devoured Bertolt, returning to human form and gaining the Colossal Titan. Meanwhile, Zeke and Reiner, the original form of the Beast Titan defeated by Levi, were rescued by friendly Titans. Before leaving, Zeke met with Eren and told Eren, "We've all been brainwashed by your father." Although Wall Maria was successfully reclaimed, only nine members of the Survey Corps survived the battle.

Eren and the Legion finally entered the basement left by Grisha Yeager, learning about Grisha Yeager's past and the truth of the world recorded therein. Eren, Mikasa, and Armin Arlert, upon their return, were imprisoned for violating military discipline. During their confinement, Eren awakened more of his father's memories, learning of many more brutal aspects of the past. During his encounter with Historia at the medal ceremony, he saw fragments of his future memories, learning that he would initiate the "War for Heaven and Earth," Rumbling and slaughtering 80% of the world's life, ultimately being stopped by his comrades.

A year later, the Titans inside and outside the walls were finally wiped out. Eren and the others were able to reach the seaside they had always dreamed of, but they learned that the battle was far from over, and the other side of the sea was full of enemies. Eren wondered if they could gain their freedom if they killed all the enemies.

Shortly after Eren and his group arrived at the seaside, Marley sent its first reconnaissance fleet to Paradis Island, which was easily defeated by the two Titans, Eren Armin Arlert. A group of Marley naval soldiers, led by a female soldier named Yelena, suddenly defected. In fact, they were under the command of Zeke Yeager and were liaisons sent to the island by Zeke to carry out his plan to liberate the Eldians.

When Yelena and others proposed Zeke's cooperation to the island's leadership, Eren confirmed it and confessed some previously concealed information: Without royal blood, if he could make contact with a Titan transformed from a royal bloodline holder, he could use the power of the Founding Titan within him to awaken a large number of Colossal Titan dormant within the three walls as a weapon—the "Rumbling," possessing the power to flatten the world. This was Paradis Island's trump card against the world's hostility, despite its technology being decades to centuries behind the rest of the world.

However, Eren had previously concealed this information because he didn't want Historia to be forced to become a Titan. Yelena and her "Anti-Marleyan Volunteers" brought advanced technology from the outside world to Paradis Island and assisted in building modern facilities such as trains and ports necessary for communication with the outside world. The first foreign guest received after the port was completed was Ms. Kiyomi Azumabito (also transliterated Kiyomi Azumabito Azmabit), a woman from the Eastern Ocean from the Hizuru, who had been on friendly terms with the Eldian Empire for generations. Kiyomi Azumabito offered to help Paradis Island face the outside world, but what she sought was the right to develop the Iceburst Stone resources on Paradis Island, while also searching for the inheritor of the bloodline of the Hizuru on the island— Mikasa.

Three years later, Eren, along with the Survey Corps members, and with the help of Yelena and others, and Kiyomi Azumabito's financial support, managed to cross the sea and visit the Marley Continent while concealing their identities. Eren encountered Ramzi, a boy he had once saved but who was about to be killed by the Rumbling he unleashed. He wanted to resist his future memories and not save him, but he still couldn't help but save him, thus realizing that the future he saw was destined to be unchangeable. Eren had spent half his life pursuing freedom, suddenly discovered that everything was predetermined from the beginning, and that he was born to be the kind of unfree livestock he had always hated the most, and he broke down and cried bitterly.

As the legion's horizons broadened, a harsh reality was also laid bare before them: even those willing to speak out for the human rights of the Subjects of Ymir outside the island regarded the Paradis Island islanders as demons to be eradicated. The world's hatred towards Paradis Island was irreparable.

Having determined that his future was already predetermined, Eren left the group alone.

Chaos in Marley

After separating from the Survey Corps, Eren adopted the alias "Kruger" and infiltrated the Eldian internment camp in Reberio, Marley, disguised as an Eldian soldier with amnesia and a missing leg. There, he met the kind-hearted young Falco Greaves, encouraging him during his warrior candidate training and earning Greaves' respect. Falco then sent several letters for Eren, conveying his plans to the Survey Corps and attempting to contact his half-brother Zeke. During this time, he also encountered an old doctor—Grisha's father and his own grandfather—who still used the alias "Kruger" to speak with him, but ultimately, the grandfather and grandson did not recognize each other.

Later, when the two brothers met, Eren pretended to accept Zeke's "Eldians Euthanasia Plan." On the night of the speech by Willy Tybal, the de facto ruler of Marley and the owner of the War Hammer Titan family, who gathered dignitaries from around the world to call for a joint effort to wipe out Paradis Island, Eren had Falco invite Reiner to the basement of a residential building next to the speech venue for a heart-to-heart talk.

Tormented by guilt and pressure, Reiner knelt before Eren, confessing and begging Eren to kill him. Eren seemed inclined to reconcile and forgive, but listening to Tyrone's impassioned speech outside the window, accusing Eren Yeager of being the source of chaos, Eren decided to fight to the end. So Eren transformed into a Titan, wreaked havoc in the speech hall, and killed Willy and many other dignitaries and civilians present.