Vol. 4War and the Tremor Fruit
Nereus and his allies challenged the World Government, fought through the Summit War, and prepared for threats across the seas. The stretch ended with Ace, Marco, Nanamo, Yamato, Blackbeard, Buckin, Weevil, and Ochoku drawn into a conflict over the Tremor-Tremor Fruit and Whitebeard’s missing body.
- Ginny and Kuma stole the Op-Op Fruit, and Nanamo organized Dressrosa’s defense against Doflamingo.
- Nanamo and her allies defeated the Donquixote Family, while Rosinante’s ties to the Marines and Vergo’s betrayal shaped the aftermath.
- Nereus and his allies expanded their influence as Chopper, Ace, and the Straw Hat Crew set out on new paths.
- Vivi sought help against Crocodile, while Jinbe and Yamato considered intervening and Ace pursued Blackbeard.
- Nereus’s conflict with the World Government widened into the Summit War, where Luffy and the Whitebeard Pirates fought to save Ace.
- After the war, Zephyr sought Nereus’s help with escaped prisoners as the Straw Hat Crew recovered on Fish-Man Island.
- Nereus pursued threats to Fish-Man Island, built alliances, and prepared Pluto while the Straw Hats trained and Uta performed on Gran Tesoro.
- Yamato seized the Tremor-Tremor Fruit as Blackbeard, Ace, Marco, Nanamo, Buckin, Weevil, and Ochoku converged over Whitebeard’s body and burial treasures.