Perona

Biodata

Feature	Information
Name	Perona (佩罗娜)
Alias(es)	Ghost Queen; Ghost Princess
Gender	Female
Affiliation	Uchiha Yi's retainer group; Thunder Emperor Legion; Divine Palace
Occupation/Role	Combatant; commander; manager of the Divine Palace
Status	Active
First Appearance	Chapter 56

Power Progression

Stage / Realm / Level	Chapter	Notes
Boundary-Breaking Haki	86	Possesses inherited Haki from Uchiha Yi; her Armament Haki suppresses Marco's Phoenix abilities in a manner compared to Seastone.
Conqueror's Haki displayed	92	Releases a battlefield-wide burst that incapacitates pirates, Marines, and weaker Vice Admirals.
Boundary-Breaking Armament Haki mastery	99	Refines her control while overwhelming Kaido; can imbue ghost clones with Haki and detonate them.
Awakened dual Devil Fruits	100	Absorbs the Soul-Soul Fruit, adding it to her Ghost-Ghost Fruit and enabling powerful physical ghost clones.
World-Breaking Haki	106	All three Haki types are described as fully developed at the World-Breaking level during her battle with Dragon.
Boundary-Breaking Ghost-Ghost and Soul-Soul Fruits	113	After ten years of training in Tsukuyomi: Illusion Realm, both Fruits reach Boundary-Breaking level and are awakened.
Mid-stage Realm-Breaking combat output	117	Can dominate Five Elders but is outmatched by the high-stage Realm-Breaking Three Saints.

Background / History

Perona first appears rebuking a man who ignored her warning to flee if danger arose. Though she mocks him, she reveals that she had followed him cautiously and likely saved his life. 56

Before the Marineford conflict, Perona had already seized financial control of the Donquixote Pirates, leaving Doflamingo dependent on her goodwill and fearful of provoking her. 71

While operating independently at sea, she tracked Kaido to Totto Land and discovered his meeting with Charlotte Linlin. Perona wanted to attack Whole Cake Island immediately because she sensed that Linlin's Soul-Soul Fruit would greatly strengthen her ghost-based abilities. Uchiha Yi deferred the attack until after Marineford but promised the Fruit to her. 80

At Marineford, Perona led Raitei's forces in blocking the Whitebeard Pirates from interfering in Uchiha Yi's fight with Whitebeard. She defeated Marco, pursued the fleeing Blackbeard Pirates, and nearly annihilated the crew before Uchiha Yi arrived to kill Blackbeard. 86 94

After Linlin's death, Perona received the Soul-Soul Fruit as promised. Its fusion with her Ghost-Ghost Fruit allowed her to create multiple powerful physical ghost clones. 99 100

Following the conquest of the One Piece World, Perona became responsible for the Divine Palace. Uchiha Yi routinely leaves major decisions to her, effectively making her one of the principal administrators of his rule. 126

Appearance

Perona is described as small and petite. She is commonly seen carrying a small floral parasol, which she also uses while fighting. 86 106

• Carries a little floral parasol. 86
• Her seemingly harmless appearance causes opponents such as Blackbeard to underestimate her. 92 94
• Often fights with a cold smile or a dismissive expression. 86 96

Personality

Perona is playful, sarcastic, and openly arrogant. She enjoys taunting opponents who underestimate her, particularly when she has already secured the advantage. 56 86 113

Despite her reluctance to admit it, she responds positively to Uchiha Yi's promises and guidance. Her desire to remain the strongest woman at his side motivates her to seek training and stronger Devil Fruit powers. 80 105

She is also calculating in battle. During the conflict with Dragon, she deliberately avoids killing him because Uchiha Yi needs his Devil Fruit intact for extraction. 107 Once she gains sufficient power, she sometimes prolongs battles for amusement rather than ending them immediately. 113

Abilities & Skills

Ghost-Ghost Fruit

Perona's primary Devil Fruit grants her ghost-based powers, including ghost doubles and emotion ghosts.

• Creates large numbers of ghost clones through Ghost Doppelganger. 94
• Produces emotion ghosts that disrupt an enemy's emotions and interfere with Devil Fruit control. 105
• Can enhance ghosts with Soul-Soul Fruit power, turning them into Homie-like entities. 105
• Reaches Boundary-Breaking level after training in Tsukuyomi: Illusion Realm. 113
• Her ghost clones can be coated in Armament Haki and detonated as explosives. 99

Soul-Soul Fruit

Perona obtains Charlotte Linlin's Soul-Soul Fruit after Linlin's defeat. 99 100

• Places souls into ghost clones, making them physical Homie-like constructs. 100
• Physical clones can wield roughly 80% of Perona's power. 100
• Can create seven or eight such clones simultaneously. 100
• Strengthens her emotion ghosts severalfold. 105
• Is awakened and developed to Boundary-Breaking level. 113

Haki

Perona possesses all three forms of Haki, inherited through her connection to Uchiha Yi and further developed through training. 86 106

• Armament Haki — Initially reaches Boundary-Breaking level, allowing her to suppress Devil Fruit users as though they were restrained by Seastone. 86 87
• Conqueror's Haki — Knocks out large numbers of Marines and pirates across the Marineford battlefield. 92
• Observation Haki — Reaches a level at which she can foresee and evade the critical points of an opponent's attacks. 113
• World-Breaking Haki — Matches Dragon's World-Breaking Devil Fruit power and allows her to repel his wind-and-lightning attacks. 105 106

Ghost Divine Sky Strike

A close-combat technique developed through special training under Uchiha Yi.

• Uses taijutsu in combination with her ghost powers. 105
• Shatters Dragon's hurricane-and-lightning attack. 105
• Is amplified by the Soul-Soul Fruit's enhancements. 105

Combat Capability

Perona is regarded as Uchiha Yi's strongest female combatant and ranks first among his ten retainers. 102 105

• Easily defeats Marco during the Marineford War. 86 87
• Battles Kaido directly, repeatedly breaking through his defenses with Boundary-Breaking Armament Haki. 96 99
• Leaves Kaido critically injured by detonating a Haki-infused ghost clone at close range. 99
• Suppresses Dragon while simultaneously maintaining several Homie ghost clones elsewhere on the battlefield. 105 107
• Defeats a bearded Five Elder before challenging one of the Three Saints. 115
• Cannot overcome the Ice-Fire Qilin Three Saint alone due to the gap between her mid-stage and the Saint's high-stage Realm-Breaking power. 117

Equipment / Items

• Floral parasol — Perona's usual accessory; she carries it into battle. 86
• Custom Dao Realm weapon — One of the weapons Uchiha Yi provided so that kills by his followers would be credited to his system; Perona uses such a weapon when killing a Five Elder. 115

Relationships

• Uchiha Yi — Her lord and benefactor. He trains her, grants her power, promises her the Soul-Soul Fruit, and relies on her to make major decisions for the Divine Palace. 80 99 113 126
• Doflamingo — Subordinate who fears provoking Perona after she gains control of the Donquixote Pirates' finances; later assists her in capturing Marco and pursuing the Blackbeard Pirates. 71 91 94
• Marco — Opponent at Marineford. Perona defeats and suppresses him before having Doflamingo guard him as a prisoner. 86 87 91
• Blackbeard — Perona defeats and psychologically breaks him after he flees Marineford with his crew. 94
• Kaido — Opponent at Totto Land. Perona overwhelms him with Boundary-Breaking Armament Haki and an explosive ghost clone. 96 99
• Monkey D. Dragon — Opponent during the war against the allied forces. Perona seeks to capture him alive so Uchiha Yi can extract his Devil Fruit. 105 107
• Gion — Fellow retainer and combatant. Perona and Gion divide the remaining Five Elders between themselves, with Perona taking the stronger bearded Elder. 112

Story Role / Major Arcs

Marineford War

Perona joins Uchiha Yi's force for the Marineford War, leads troops against Whitebeard's commanders, defeats Marco, and uses Conqueror's Haki to shift the wider battlefield. 80 86 92

Blackbeard Pirates Pursuit

Perona detects Blackbeard's escape, pursues his crew with Doflamingo, and kills nearly everyone except Blackbeard and Catarina Devon before Uchiha Yi arrives. 94

Totto Land Conflict

Perona's pursuit of the Soul-Soul Fruit drives the assault on Totto Land. She fights Kaido while Uchiha Yi faces Charlotte Linlin, then receives Linlin's Fruit after the battle. 80 96 99 100

War Against Dragon

Perona intercepts Dragon, counters his World-Breaking weather ability with her own World-Breaking Haki, and attempts to capture him rather than kill him. 105 106 107

Assault on Mary Geoise

Perona defeats a Five Elder, demonstrates the Boundary-Breaking development of both of her Devil Fruits, and later confronts the Ice-Fire Qilin Three Saint before Uchiha Yi intervenes. 112 113 115 117

Divine Palace Administration

After Uchiha Yi establishes the Divine Hall and becomes Holy Lord, Perona manages the Divine Palace and makes major decisions in his stead. 125 126

Trivia

• Perona initially showed no interest in the Devil Fruits distributed by Uchiha Yi, but immediately chose the Soul-Soul Fruit once she encountered it. 95
• She was the first person whose real body entered Uchiha Yi's Tsukuyomi: Illusion Realm for training. 113
• One minute in the outside world equaled ten years of training for Perona within the Illusion Realm. 113
• Blackbeard calls her inhuman after being repeatedly defeated by her. 94