Perona

Biodata

Power Progression

Background / History

• Perona is first introduced residing on Thriller Bark, where she commands the animal and wild zombies and assists Gecko Moria in capturing pirate crews and gathering shadows 80 97.
• After Thriller Bark is infiltrated and disrupted by the Straw Hat Pirates, Perona uses her Negative Ghosts to incapacitate Luffy, Zoro, Franky, and others before being thwarted by Xi Yan's illusions and Usopp's natural negativity 85 98.
• Following the events of Thriller Bark and the war, Perona spends a two-year training period alongside Roronoa Zoro, developing her Devil Fruit abilities and forming a protective, complex bond with him 339 340.
• She later reunites with the Straw Hat Pirates during the Dressrosa arc, serving as an intelligence gatherer, scout, and combat ally using her ghosts and "Ghost Communication" to protect the crew and their allies 538 559.

Appearance

Perona is a young woman characterized by a distinct, doll-like Gothic aesthetic. She typically wears ornate dresses, a crown, and heavy makeup, sporting long, voluminous hair and carrying an umbrella.

• Distinguishing Features:
  • Large, expressive eyes and distinctive crown-like headwear.
  • Gothic-styled Lolita dresses.
  • Ability to manifest a floating, semi-transparent ghost form.

Personality

Perona possesses a proud, confident, and somewhat childish demeanor, often referring to herself as "this young lady" and laughing with her signature "Haraharahara~" 85 98. She enjoys cute things and children, frequently showing a softer side when interacting with sweet-talking kids 422. Despite her dramatic and prideful attitude, she is deeply loyal to those she cares about—most notably Gecko Moria, whom she worries about like a father, and Zoro, whom she aided and trained alongside for two years 339 559.

Abilities & Skills

Hollow-Hollow / Spirit-Spirit Fruit

Perona consumed a Paramecia-class Devil Fruit that grants her mastery over spirits and ghosts, allowing her to project her soul, phase through physical barriers, and manipulate various types of spectral constructs 80 339 481.

• Negative Ghost: Sends out small, semi-transparent ghosts that pass through targets, draining all their fighting spirit and plunging them into severe depression and extreme self-loathing 97 350.
• Exploding Ghost: Manifests ghosts capable of detonating with immense destructive force, bypassing physical durability unless countered by Haki 423 481.
• Ghost Communication: Uses detached ghost clones to scout vast distances and transmit real-time visual and auditory information back to her main body 559.
• Flight: Capable of levitating and flying freely through the air at high speeds 384 426.

Equipment / Weapons / Items

• Umbrella: Often carried as part of her Gothic attire.
• Den Den Mushi: Utilized for long-distance communication alongside her ghost projection abilities 697.

Relationships

• Gecko Moria — Father figure and former captain; Perona deeply cares for his well-being and seeks to find him in Wano Country 97 486.
• Roronoa Zoro — Savior and companion; they spent two years together training on Kuraigana Island, developing a close, bantering dynamic 339 340.
• Xi Yan — Acquaintance and occasional adversary; Perona is deeply intimidated by his Sharingan and illusion techniques, which successfully countered her abilities 85 98.
• Usopp — Former enemy; Usopp became completely immune to her Negative Ghosts due to his naturally pessimistic personality 98.
• Nami — Ally; Perona temporarily swapped bodies with her during the Punk Hazard arc, leading to mutual cooperation and shared adventures 426 438.
• Merry — Ally and ship spirit; Perona frequently interacts and works alongside her during missions 351 562.

Story Role / Major Arcs

• Thriller Bark Arc: Acts as a primary antagonist under Moria, deploying zombie forces and Negative Ghosts against the invading Straw Hat Pirates before being defeated by Xi Yan's genjutsu 80 85.
• Two-Year Timeskip: Resides on Kuraigana Island alongside Mihawk and Zoro, where she refines her Devil Fruit capabilities to turn her entire physical body into spirit form 339.
• Dressrosa Arc: Joins forces with the Straw Hat alliance to undermine Doflamingo's regime, utilizing her scouting ghosts and psychological warfare to protect the crew and assist the Tontatta tribe 538 558.

Notable Quotes

"Lord Moria, another group of pirates has triggered our trap." 80

"Now you know how amazing this young lady is~" 350

"Who... who wants to hide behind this idiot? I am the Ghost Princess, and I will definitely learn Haki!" 559